Ann Dowd is an American actress. She had her career beginnings in the 1980s, starting on television. Later in 1990, she made her big-screen debut in the film Green Card. Soon after, she had roles in many more films, including Philadelphia and It Could Happen to You. She next starred as Louise Preston in the movie Shiloh and its sequel Shiloh 2: Shiloh Season. On TV, her first starring role was as Sister Maureen “Mo” Brody in the TV drama Nothing Sacred. In the 2000s, she had roles in several movies, including Garden State, The Manchurian Candidate, The Notorious Bettie Page, Flags of Our Fathers, and Marley & Me. Later in 2013, she played Estabrook Masters in Masters of Sex. After this, she starred as Patti Levin in The Leftovers. As of 2017, she stars as Aunt Lydia Clements in the Hulu series The Handmaid’s Tale.